1. South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, PR China
2. State Key Laboratory of Applied Microbiology Southern China, Guangzhou 510070, PR China
3. University of Chinese Academy of Sciences, Beijing 100049, PR China
4. Guangdong Provincial Key Laboratory of Microbial Culture Collection and Application, Guangdong Institute of Microbiology, Guangzhou 510070, PR China