1. 1School of Natural and Environmental Sciences, Newcastle University, Devonshire Building, Newcastle upon Tyne NE1 7RU, UK
2. 2Centre for Biotechnology and Bioengineering, University of Chile, Beauchef 851, Santiago, Chile
3. 3Laboratory of Microbial Complexity and Functional Ecology, Departamento de Biotecnología, Facultad de Ciencias del Mary Recursos Biológicos and Centre for Biotechnology and Bioengineering, Universidad de Antofagasta, Chile
4. 4Leibniz Institute DSMZ-German Collection of Microorganisms and Cell Cultures, Germany