1. University of Chinese Academy of Sciences, 100049, Beijing, PR China
2. Institute of Oceanology, Chinese Academy of Sciences, 266071, Qingdao, PR China
3. Department of Bioengineering, College of Marine Sciences and Biological Engineering, Qingdao University of Science and Technology, 266042, PR China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences, 7 Nanhai Road, Qingdao, 266071, PR China