1. School of Life Sciences, Xiamen University, Xiamen, Fujian, 361102, PR China
2. Key Laboratory of the Ministry of Education for Coastal and Wetland Ecosystems, Xiamen University, Xiamen, Fujian, 361102, PR China
3. Key Laboratory of Marine Biogenetic Resources, Third Institute of Oceanography, State Oceanic Administration, Xiamen, Fujian, 361102, PR China