1. Department of Biological Sciences, Tokyo Metropolitan University, 1-1, Minami-Osawa, Hachioji, Tokyo, 192-0397, Japan
2. Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), 1–1–1, Higashi, Tsukuba, Ibaraki, 305–8566, Japan
3. NITE Biological Resource Center (NBRC), National Institute of Technology and Evaluation (NITE), 2–5–8, Kazusakamatari, Kisarazu, Chiba, 292–0818, Japan
4. Department of Molecular Microbiology, Tokyo University of Agriculture, 1-1-1, Sakuragaoka, Setagaya-ku, Tokyo, 156-8502, Japan