1. 1Department of Microbial, Biochemical and Food Biotechnology, University of the Free State, Bloemfontein, Free State, South Africa
2. 2Department of Food Science and Technology, Cape Peninsula University of Technology, Bellville, Western Cape, South Africa
3. 3Department of Biotechnology, Institute for Microbial Biotechnology and Metagenomics, University of the Western Cape, Bellville, Western Cape, South Africa
4. 4Biology Department, Lycoming College, Williamsport PA 17701, USA