1. College of Chemistry and Biological Engineering, University of Science and Technology Beijing, Beijing 100083, PR China
2. University of Science and Technology (UST), Daejeon 34113, Republic of Korea
3. Biological Resource Center/Korean Collection for Type Cultures (KCTC), Korea Research Institute of Bioscience and Biotechnology, Jeonbuk 56212, Republic of Korea
4. Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing 100081, PR China