1. 1Atmosphere and Ocean Research Institute, The University of Tokyo, Kashiwa, Chiba, Japan
2. 2Division of Biological Sciences, College of Arts and Sciences, University of the Philippines Visayas, Iloilo, Philippines
3. 3Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Yokosuka, Japan
4. 4Department of Bacteriology, Faculty of Medical Sciences, Kyushu University, Fukuoka, Japan