1. Centre for Research in Bioscience, Faculty of Health and Life Sciences, University of the West of England, Frenchay Campus, Bristol, UK
2. Japan Collection of Microorganisms (RIKEN-BRC JCM), RIKEN Bioresource Centre, Tsukuba, Ibaraki 305-0074, Japan
3. Forest Research, Centre for Ecosystems, Society and Biosecurity, Alice Holt Lodge, Surrey, UK