1. State Key Laboratory of Microbial Technology, Shandong University, Qingdao 266237, PR China
2. School of Life Sciences and Medicine, Shandong University of Technology, Zibo 255000, PR China
3. School of Pharmacy, Binzhou Medical University, Yantai 264003, PR China
4. Laboratory for Marine Biology and Biotechnology, Qingdao Marine Science and Technology Center, Qingdao 266237, PR China