1. College of Marine Science, Shandong University at Weihai, Weihai 264209, PR China
2. Crop Bioprotection Research Unit, National Center for Agricultural Utilization Research, Agricultural Research Service, U.S. Department of Agriculture, Peoria, IL 61604, USA
3. State Key Laboratory of Microbial Technology, Shandong University, Jinan 250100, PR China