1. National Research Institute of Aquaculture,Fisheries Research Agency, 422-1 Nakatsuhamaura, Minami-ise, Mie 516-0193,Japan
2. National Research Institute of Fisheries Science,Fisheries Research Agency, 2-12-4 Fukuura, Kanazawa, Yokohama, Kanagawa, 236-8648,Japan
3. Laboratory of Genome Science,Tokyo University of Marine Science and Technology, 4-5-7 Konan, Minato, Tokyo, 108-8477,Japan
4. Fisheries Research Division,Oita Prefectural Agriculture, Forestry and Fisheries Research Center, 194-6 Tsuiura, Kamiura, Saiki, Oita, 879-2602,Japan