1. Center for Ocean Mega-Science, Chinese Academy of Sciences, 7 Nanhai Road, Qingdao 266071, PR China
2. Center of Deep Sea Research, Institute of Oceanology, Chinese Academy of Sciences, 7 Nanhai Road, Qingdao 266071, PR China
3. Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, PR China
4. University of Chinese Academy of Sciences, Beijing 100049, PR China
5. Key Laboratory of Marine Ecology and Environmental Sciences, Chinese Academy of Sciences, Qingdao 266071, PR China