1. School of Life Science, Liaoning Normal University, Dalian 116029, PR China
2. Dandong Forestry and Grassland Development Service Center, Dandong 118000, PR China
3. State Key Laboratory of Biocontrol and Guangdong Provincial Key Laboratory of Plant Resources, School of Life Science, Sun Yat-Sen University, Guangzhou, 510275, PR China
4. CAS Key Laboratory of Biogeography and Bioresource in Arid Land, Xinjiang Institute of Ecology and Geography, Urumqi, 830011, PR China