1. College of Resources and Environment, Tibet Agricultural and Animal Husbandry University, Linzhi 860000, PR China
2. CAAS-CIAT Joint Laboratory in Advanced Technologies for Sustainable Agriculture, Institute of Agricultural Resources and Regional Planning, Chinese Academy of Agricultural Sciences, Beijing 100081, PR China
3. Jining Academy of Agricultural Sciences, Jining 272000, PR China
4. Key Laboratory of Microbial Resources Exploitation and Application of Gansu Province, Institute of Biology, Gansu Academy of Sciences, Lanzhou 730000, PR China
5. College of Life Sciences, Yantai University, Yantai 264005, PR China