1. Université de Lyon, F-69622, Lyon, France; Université Lyon 1, Villeurbanne, France; CNRS, UMR5557, Ecologie Microbienne, Villeurbanne, France; INRA, UMR1418, Villeurbanne, France
2. Department of Ecology, Environment and Plant Sciences, Stockholm University, 106 91 Stockholm, Sweden
3. Department of Plant Sciences, University of California, Davis, CA 95616, USA
4. Department of Biology, Colorado State University, Pueblo, CO 81001, USA