1. Coral Reef Research Center of China, Guangxi University, Nanning 530004, PR China
2. School of Marine Sciences, Guangxi University, Nanning 530004, PR China
3. Key Laboratory of Tropical Marine Bio-resources and Ecology (LMB), Guangdong Key Laboratory of Marine Materia Medica (LMMM-GD), South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, PR China
4. Guangxi Key Laboratory on the Study of Coral Reefs in the South China Sea, Nanning 530004, PR China