1. Department of Microbiology, Faculty of Science, Chulalongkorn University, 254 Phayathai Road, Pathumwan, Bangkok 10330, Thailand
2. Microbe Division/Japan Collection of Microorganisms (JCM), RIKEN BioResource Center, 3-1-1 Koyadai, Tsukuba, Ibaraki, Japan 305-0074
3. Department of Biochemistry and Microbiology, Faculty of Pharmaceutical Sciences, Chulalongkorn University, 254 Phayathai Road, Pathumwan, Bangkok 10330, Thailand