1. State Key Laboratory of Lake Science and Environment, Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences, Nanjing 210008, PR China
2. University of Chinese Academy of Sciences, Beijing 100049, PR China
3. Tibet Plateau Institute of Biology, Lhasa 850000, PR China
4. Sino-Danish Centre for Education and Research, University of Chinese Academy of Sciences, Beijing 100049, PR China