1. 1Tibet Plateau Institute of Biology, Lhasa 850000, PR China
2. 2State Key Laboratory of Lake Science and Environment, Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences, Nanjing 210008, PR China
3. 3University of Chinese Academy of Sciences, Beijing 100049, PR China
4. 4College of Life Sciences, Nanjing Normal University, Nanjing 210046, PR China