1. Tibet Plateau Institute of Biology, Lhasa 850000, PR China
2. College of Life Sciences, Nanjing Normal University, Nanjing 210046, PR China
3. State Key Laboratory of Lake Science and Environment, Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences, Nanjing 210008, PR China
4. University of Chinese Academy of Sciences, Beijing 100049, PR China
5. College of Life Sciences, Anhui Normal University, Wuhu 241002, PR China
6. Sino-Danish Centre for Education and Research, University of Chinese Academy of Sciences, Beijing 100049, PR China