1. Department of Biology, Jeju National University, Jeju 690-756, Republic of Korea
2. Jeju Biological Resource Co., Ltd, CTC Business Incubator Jeju Tourism College, Jeju 690-756, Republic of Korea
3. Research Institute for Basic Science, Jeju National University, Jeju 690-756, Republic of Korea
4. Department of Aquatic Life Medicine, Jeju National University, Jeju 690-756, Republic of Korea