1. R&D Center of Marine Biotechnology, Institute of Oceanology, Chinese Academy of Sciences,Qingdao 266071,PR China
2. State Key Laboratory of Microbial Resources, Institute of Microbiology, Chinese Academy of Sciences,Beijing 100101,PR China
3. Nantong R&D Center of Marine Science and Technology, Institute of Oceanology, Chinese Academy of Sciences,Nantong 226019,PR China