1. Department of Biotechnology and Food Technology, Faculty of Science, University of Johannesburg, Doornfontein Campus, PO Box 17011, Doornfontein, 2028, Johannesburg, South Africa
2. Department of Biochemistry, Faculty of Science, University of Johannesburg, Kingsway Campus, PO Box 524, Auckland Park, 2006, Johannesburg, South Africa
3. Key Laboratory of Agricultural Microbiomics and Precision Application (MARA), State Key Laboratory of Applied Microbiology Southern China,, Institute of Microbiology, Guangdong Academy of Sciences, Guangzhou 510070, People’s Republic of China