1. Department of Veterinary Disease Biology, Faculty of Health and Medical Sciences, University of Copenhagen, Stigboejlen 4, 1870 Frederiksberg C, Denmark
2. Center for Zoo and Wild Animal Health, Copenhagen Zoo, 2000 Frederiksberg, Denmark
3. Department of Bioscience, Faculty of Science and Technology, Arctic Research Centre, Frederiksborgvej 399, PO Box 358, Aarhus University, 4000 Roskilde, Denmark