1. University of Liverpool, Institute of Infection and Global Health, Leahurst Campus, Neston, CH64 7TE, UK
2. Unidad de Epidemiología y Medicina Preventiva, Facultad de Veterinaria, Universidad de Las Palmas de Gran Canaria, C/Trasmontaña s/n, Arucas, 35413, Islas Canarias, Spain
3. The Oaks, Nutshell Lane, Upper Hale, Farnham, Surrey, GU9 0HG, UK
4. University of Liverpool, Institute of Integrative Biology, Liverpool, L69 7ZB, UK