1. 2University of Chinese Academy of Sciences, Beijing 100049, PR China
2. 1Key Laboratory of Biofuels, Shandong Provincial Key Laboratory of Energy Genetics, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao 266101, PR China
3. 3Institute of Marine Science and Technology, Shandong University, Qingdao 266273, PR China
4. 4Tianjin Branch of China National Offshore Oil Corporation Ltd, Tianjin 300450, PR China