1. Joint Lab for Ocean Research and Education at Dalhousie University, Shandong University and Xiamen University, Qingdao 266237, PR China
2. Institute of Marine Science and Technology, Shandong University, Qingdao 266237, PR China
3. Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519000, PR China
4. Marine College, Shandong University, Weihai 264209, PR China