1. Aix Marseille Univ, IRD, APHM, MEPHI, 13005 Marseille, France
2. IHU Méditerranée Infection, 13005 Marseille, France
3. Aix Marseille Univ, IRD, APHM, VITROME, 13005 Marseille, France
4. Faculty of Science, M’Hamed Bougara Boumerdès University, Boumerdès 35000, Algeria
5. VITROME, Campus International IRD-UCAD Hann, Dakar, Senegal