1. Japan Collection of Microorganisms, RIKEN BioResource Research Center, Tsukuba, Ibaraki 305-0074, Japan
2. Graduate School of Life Sciences, Toyo University, Oura, Gunma 374-0193, Japan
3. Present address: Faculty of Science and Engineering, Soka University, Tokyo 192-8577, Hachioji, Japan
4. Research Institute for Geo-Resources and Environment, National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, 305-8567, Japan