1. Graduate School of Agricultural Science, Tokyo University of Agriculture and Technology, Fuchu, Tokyo 183-8509, Japan
2. Department of Forest Science, Faculty of Regional Environmental Science, Tokyo University of Agriculture, Setagaya,Tokyo 156-8502, Japan
3. Institute of Global Innovation Research, Tokyo University of Agriculture and Technology, Fuchu, Tokyo 183-8509, Japan
4. Illinois Natural History Survey, University of Illinois, 1816 S. Oak Street, Champaign, IL 61820, U.S.A.
5. Central Region Agricultural Research Center, National Agriculture and Food Research Organization (NARO), Tsukuba, Ibaraki, Japan