1. Field Science Center, Hokkaido University, Hirai 559, Kozagawa, Wakayama 649-4563, Japan
2. Department of Biology, Faculty of Science, West 1-C-915, Kyushu University, Moto-oka 744, Fukuoka 819-0395, Japan
3. Graduate School of Systems Life Sciences, West 1-C-804, Kyushu University, Moto-oka 744, Fukuoka 819-0395, Japan
4. Field Science Center, Hokkaido University, North 11, West 10, Sapporo 060-0811, Japan