1. Fisheries Resources Institute, Japan Fisheries Research and Education Agency, 116 Katsurakoi, Kushiro, Hokkaido 085-0802, Japan
2. Department of Marine Biosciences, Tokyo University of Marine Science and Technology, 4-5-7, Konan, Minato-ku, Tokyo 108-8477, Japan
3. Division of Marine Bio-resource and Environmental Science, Graduate School of Fisheries Sciences, Hokkaido University, 3-1-1, Minato-cho, Hakodate, Hokkaido 041-8611, Japan