1. Kyoto University Museum, Kyoto University, Yoshida-honmachi, Sakyo, Kyoto 606-8501, Japan
2. Laboratory of Animal Genetics and Breeding, Graduate School of Bioagricultural Sciences, Nagoya University, Furo-Cho, Chikusa, Nagoya 464-8601, Japan
3. Department of Livestock and Aquaculture Research, Nay Pyi Taw, 15013, Myanmar
4. Department of Zoology, Yangon University of Distance Education, Yangon, 11041, Myanmar
5. Institute of Low Temperature Science, Hokkaido University, Kita-ku, Sapporo 060-0819, Japan