1. Faculty of Science, Toho University, Funabashi, Chiba 274-8510, Japan
2. Division of Agriculture, Forestry and Fisheries, Oita Prefectural Office, Ote-machi, Oita, Oita 870-8501, Japan
3. Faculty of Environmental Science, University of human Environments, Okazaki, Aichi 444-3505, Japan
4. Graduate School of Fisheries and Environmental Sciences, Nagasaki University, Bunkyo, Nagasaki 852-8521, Japan
5. Graduate School of Frontier Sciences, The University of Tokyo, Kashiwanoha, Kashiwa, Chiba 277-8563, Japan
6. Faculty of Fisheries, Kagoshima University, Shimoarata, Kagoshima 890-0056, Japan