1. Division of Bioscience and Bioindustry, Graduate School of Technology, Industrial and Social Sciences, Tokushima University, 2-1 Minami-josanjima, Tokushima 770-8513, Japan
2. Shikoku Institute of Natural History, 470-1 Shimobun-otsu, Susaki 785-0023, Japan
3. Botanical Gardens, Tohoku University, Kawauchi 12-2, Aoba, Sendai 980-0862, Japan