1. United Graduate School of Agricultural Science, Tokyo University of Agriculture and Technology, 35-8 Saiwai, Fuchu 183-8509, Japan
2. Department of Ecology, UNESP São Paulo State University, Botucatu, São Paulo 18610-034, Brazil
3. Institute of Agriculture, Tokyo University of Agriculture and Technology, 3-5-8 Saiwai, Fuchu 183-8509, Japan