1. Graduate School of Environmental Science, Hokkaido University, Sapporo, Hokkaido, Japan
2. Laboratory of Bioresources, Applied Biology Co. Ltd, Minato-ku, Tokyo, Japan
3. Forestry Research Institute, Hokkaido Research Organization, Bibai, Hokkaido, Japan
4. Graduate School of Life Science, Hokkaido University, Sapporo, Hokkaido, Japan