1. Department of Biology, Faculty of Sciences, Kyushu University, Fukuoka 812-8581, Japan.
2. Graduate School of Fisheries and Environmental Studies, Nagasaki University, 1–14 Bunkyo-machi, Nagasaki 852-8521, Japan.
3. Research Institute for the Environment and Livelihoods, Charles Darwin University, Darwin, NT 0909, Australia.