1. Centro Nacional Patagónico, Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Boulevard Brown 2915, U9120ACF, Puerto Madryn, Chubut, Argentina.
2. University of Washington, Department of Biology, Box 351800, Seattle, WA 98195-1800, USA.
3. Centre for Wildlife Ecology, Simon Fraser University, 5421 Robertson Road, Delta, BC, V4K 3N2, Canada.