1. CIBIO-InBIO, Research Center in Biodiversity and Genetic Resources, InBIO Associate Laboratory, Campus de Vairão, 7 Rua Padre Armando Quintas, 4485-661 Vairão, Portugal.
2. CEFE, Université de Montpellier, CNRS, Montpellier, France.
3. Department of Biology, San Francisco State University, 1600 Holloway Avenue, San Francisco, CA 94132, USA.