1. The Kennel Club, Clarges Street, London W1J 8AB, United Kingdom.
2. School of Veterinary Medicine and Science, University of Nottingham, Sutton Bonington Campus, Loughborough LE12 5NT, United Kingdom.
3. Animal Breeding & Genomics Centre, Wageningen UR Livestock Research and Centre for Genetic Resources, The Netherlands, Wageningen University & Research, P.O Box 338, 6700 AH Wageningen, the Netherlands.