1. Institut de l’Elevage, UMT eBIS, 78350 Jouy-en-Josas, France.
2. Université Paris Saclay, INRAE, AgroParisTech, GABI, Domaine de Vilvert Bat 211, 78350 Jouy-en-Josas, France.
3. Wageningen University and Research, Animal Breeding and Genomics, P.O. Box 338, 6700 AH Wageningen, the Netherlands.