1. Department of Plant Biology, University of Illinois at Urbana-Champaign, Urbana, Illinois 61801, USA;
2. Smithsonian Tropical Research Institute, Apartado 0843-03092, Panamá, República de Panamá
3. Department of Crop Sciences, University of Illinois at Urbana-Champaign, Urbana, Illinois 61801, USA;
4. School of Plant Sciences and Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, Arizona 85721, USA;
5. Department of Integrative Biology, University of South Florida, Tampa, Florida 33620, USA;,