1. Departments of Entomology and Integrative Biology, Michigan State University, East Lansing, Michigan, USA
2. Current affiliation: Department of Land Resources and Environmental Sciences, Montana State University, Bozeman, Montana, USA; email:
3. Department of Biological Science, Florida State University, Tallahassee, Florida, USA
4. Entomology and Nematology Department, University of Florida, Gainesville, Florida, USA
5. Department of Biological Sciences, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, USA