1. Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, Arizona, 85721-0088;
2. Zoologisches Museum, Universität Zürich, CH-8057 Zürich, Switzerland;
3. Department of Zoology, Institute of Ecology and Earth Sciences, University of Tartu, 51014 Tartu, Estonia;
4. Department of Entomology, University of Kentucky, Lexington, Kentucky, 40546-0091;