1. Department of Biomedical Sciences, Cornell University, Ithaca, New York 14853;
2. Department of Zoology, University of Cambridge, Cambridge CB2 3EJ, United Kingdom;
3. Integrative and Systems Biology Research Theme, Anderson College Complex, University of Glasgow, Glasgow G11 6NU, United Kingdom, and College of Clinical Laboratory Sciences, King Saud University, Riyadh, Saudi Arabia;