1. School of Environmental and Rural Science, University of New England, Armidale, New South Wales 2351, Australia;
2. Fenner School of Environment and Society, College of Science, The Australian National University, Canberra ACT 2601, Australia
3. The New Zealand Institute for Plant and Food Research Limited, Christchurch 8140, New Zealand
4. Department of Biology, University of Maryland, College Park, Maryland 20742, USA
5. Rocky Mountain Biological Laboratory, Crested Butte, Colorado 81224, USA