1. Department of Ecology, Evolution and Behavior, University of Minnesota, St. Paul, Minnesota 55108;
2. Doñana Biological Station, Spanish Research Council CSIC, C/Americo Vespucio s/n, 41092, Isla de la Cartuja, Seville, Spain;
3. Center for Evolutionary Biology, School of Animal Biology, The University of Western Australia, Crawley, Washington 6009, Australia;,
4. Department of Biological Sciences, Macquarie University, NSW 2109, Australia;