1. CSIRO Ecosystem Sciences and Sustainable Agriculture Flagship, Canberra, ACT 2601, Australia;
2. University of Washington, Interdisciplinary Arts and Sciences, Tacoma, Washington 98402;
3. Washington State University, Puyallup Research & Extension Center, Puyallup, Washington 98371;
4. CSIRO Ecosystem Sciences, Narrabri, NSW 2390, Australia;